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The ensemble and supporting cast feature Rashida Jones as Ann Perkins, Paul Schneider as Mark Brendanawicz, Aziz Ansari as Tom Haverford, Nick Offerman as Ron Swanson, Aubrey Plaza as April Ludgate, Chris Pratt as Andy Dwyer, Adam Scott as Ben Wyatt, Rob Lowe as Chris Traeger, Jim O'Heir as Garry "Jerry", "Larry", or "Terry" Gergich, Retta as Donna Meagle, and Billy Eichner as Craig Middlebrooks.

Poehler's character, Leslie Knope, underwent major changes after the first season, in response to audience feedback that she seemed unintelligent and "ditzy".

K., Paul Rudd, Henry Winkler, and Jon Hamm, have been featured in the series, and their characters often appear in multiple episodes.

Parks and Recreation was part of NBC's "Comedy Night Done Right" programming during its Thursday night prime-time block.

Meanwhile, Andy develops feelings for April, but she fears he is still hung up on Ann.

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